'Lionesses' in the Bible

An oracle about the animals of the Negev: "Through a land of trouble, dryness, and distress, of lionesses and roaring lions, where there is no water, a land of vipers and darting snakes, he carries their riches on donkeys' backs, and their treasures on the humps of camels, to a nation that cannot benefit them,

Where is the habitation of lionesses? And a feeding-place it is for young lions Where walked hath a lion, an old lion, A lion's whelp, and there is none troubling.

The lion did tear in pieces enough for his whelps, and strangled for his lionesses, and filled his holes with prey, and his dens with ravin.